"This is a historic day in the struggle for animal rights and in defence of our evolutionary comrades, which will doubtless go down in the history of humanity," said Pedro Pozas, Spanish director of the Great Apes Project.

Spain may be better known abroad for bull-fighting than animal rights but the new measures are the latest move turning once-conservative Spain into a liberal trailblazer.

Spain did not legalise divorce until the 1980s, but Prime Minister Jose Luis Rodriguez Zapatero's Socialist government has legalised gay marriage, reduced the influence of the Catholic Church in education and set up an Equality Ministry.

The new resolutions have cross-party or majority support and are expected to become law and the government is now committed to update the statute book within a year to outlaw harmful experiments on apes in Spain.

"We have no knowledge of great apes being used in experiments in Spain, but there is currently no law preventing that from happening," Pozas said.

Keeping apes for circuses, television commercials or filming will also be forbidden and breaking the new laws will become an offence under Spain's penal code.

Keeping an estimated 315 apes in Spanish zoos will not be illegal, but supporters of the bill say conditions will need to improve drastically in 70 percent of establishments to comply with the new law.

Philosophers Peter Singer and Paola Cavalieri founded the Great Ape Project in 1993, arguing that "non-human hominids" like chimpanzees, gorillas, orang-utans and bonobos should enjoy the right to life, freedom and not to be tortured. (Reporting by Martin Roberts; Editing by Richard Williams)